Tag: prehistoric animal

  • Indricotherium: The Largest Mammal Ever to Walk the Earth

    Indricotherium: The Largest Mammal Ever to Walk the Earth

    • Indricotherium, an enormous herbivorous mammal, lived in Asia.
    • Resembling rhinos, they lacked horns and had long necks.
    • Names like Indricotherium, Paraceratherium, and Baluchitherium are often used interchangeably to refer to these animals.

    Extinct in the 30–20-million-year-old Middle Oligocene–Early Miocene epochs, Indricotherium (a Latin word from “indrik“—a unicorn-like mythological beast in Russian mythology and from ancient Greek Θηρίον—animal) is a genus of mammals in the family Hyracodontidae. Most indricothere fossils have been unearthed around Asia. Beluchitherium, discovered in the Oligocene of Mongolia, and Aralotherium, discovered in more recent strata in the Aral Sea in Kazakhstan are close relatives of Indricotherium.

    Features of Indricotherium

    Indricotherium (also Paraceratherium, or Baluchitherium) hind legs.
    Indricotherium (also Paraceratherium, or Baluchitherium) hind legs. (Ryan Somma, cc by sa 2.0)

    The enormous Oligocene animals known as Indricotherium originated in Asia. It was a relative of modern rhinos but was hornless. Its diet consisted mostly of the foliage of towering trees, since it was an herbivore. Indricotherium fossils have been discovered in Central Asia as well.

    How Did Indricotherium Look Like?

    They resembled rhinoceroses in many ways, including their body forms, long and straight three-toed legs with a substantially expanded middle toe, and a small head on an unusually long neck. However, they did not have horns and had a domed skull instead of a flat one.

    The skull of a Indricotherium (or Paraceratherium) transouralicum. 1923.
    The skull of a Indricotherium (or Paraceratherium) transouralicum. 1923.

    Size and Weight

    The largest ever mammals were the Indricotherium and Beluchitherium, which could grow to be 18 feet (5.5 m) in height at the shoulder and 28 feet (8.5 m) in length. These animals weighed 20 to 25 tons. The length of Indricotherium’s skull was almost 4.6 feet (1.4 m).

    Indricotherium, (also known as Paraceratherium, and Baluchitherium)
    Indricotherium, (also known as Paraceratherium, and Baluchitherium). Candace Lindemann, cc by sa.

    Their Diet and Habitat

    The legs were long and huge, much like an elephant’s today. They ate grass and the leaves and branches of bushes and trees. Some of them lived in semiarid or desert environments, while others preferred damper environments like woods or wetlands.

    Perhaps the emergence of more specialized big herbivorous animals contributed to the demise of indricotheres (the members of Indricotherium). While giraffes and proboscideans specialized in eating tree branches, short-legged rhinoceroses were better suited to devouring low grassland plants.

    Indricotherium, Paraceratherium, and Baluchitherium

    Although the species within the genus Indricotherium are well-known, the taxonomy of the genus itself is currently in flux. The first member of the genus Paraceratherium wasn’t named until 1911, when the English paleontologist Clive Forster-Cooper did so. Baluchitherium was another organism he described in 1913. A. A. Borissiak first described the genus Indricotherium in 1915.

    Paraceratherium, Baluchitherium, and Indricotherium are all seen as interchangeable terms (Lucas & Sorbus, 1989) and hence equivalent. These names represent different genera within the family Hyracodontidae.

    However, there are many who believe the two genera, Indricotherium and Paraceratherium, are distinct. Whatever the case may be, these names all refer to creatures that are around the same size and shape.

    Indricotherium transouralicum: Middle to late Oligocene Indricotherium transouralicum (Pavlova, 1922) is the most common and well-studied species among others. Its historical range included what are now parts of Kazakhstan, Mongolia, Inner Mongolia, and northern China.

    Paraceratherium orgosensis: The biggest indricothere is Paraceratherium orgosensis (Chiu, 1973). The Xinjiang Uygur area of northwest China is where it was found. Paraceratherium lipidus (Xu and Wang, 1978), Dzungariotherium turfanensis (Chiu, 1973), and Dzungariotherium orgosensis (Chiu, 1973) are all likely to refer to the same species of fossil.

    Paraceratherium zhajremensis: The indricothere Paraceratherium zhajremensis (Osborn, 1923) was discovered in the Oligocene of India.

    Indricotherium prohorovi: The eastern Kazakh middle and late Miocene are home to the Indricotherium known as Indricotherium prohorovi (Borissiak, 1939).

    Paraceratherium bugtiense: The original species of the genus Paraceratherium is found in the middle Miocene of Pakistan and was named by Pilgrim (1908).

    Baluchitherium osborni: It is a more recent but etymologically equivalent name (Forster Cooper, 1913) to Paraceratherium bugtiense. It was discovered in Baluchistan’s Chitarwata Formation.

    Paraceratherium linxiaense: In the Linxia-Hui Autonomous District of Gansu Province, at the northeastern edge of the Tibetan Plateau, Paraceratherium linxiaense (Tao Deng, 2021) was found in late Oligocene sediments (26.57 million years old). At the time of its description, it was estimated to weigh 24 tons and stand 13 feet (4 m) tall (around 27 feet high at the shoulder).

    Indricotherium in the Cultural Realm

    • Paraceratherium or Indricotherium was the inspiration for the design of the walking battle machines AT-AT used by the Empire in the “Star Wars” film series.
    • V. A. Obruchev mentions indricothere in his book Plutonia.
    • The third episode of the British documentary series “Walking with Beasts” is devoted to the indricothere’s early years.
  • Patriofelis: The Father of Cats, A Jaguar-Like Prehistoric Animal

    Patriofelis: The Father of Cats, A Jaguar-Like Prehistoric Animal

    The fossils of the extinct Oxyaenodonta mammal Patriofelis (genus Patriofelis) have been discovered in North America, and they existed during the middle Eocene (about 48 to 40 million years ago). Although the resemblances between this predator and a modern jaguar were the consequence of convergent evolution, they were still strikingly similar. Patriofelis was a massive Oxyaenodonta (“sharp hyena”), measuring in at over 100 inches (250 cm) in length (including its tail).

    The Discovery of Patriofelis

    Patriofelis The Father of Cats
    Patriofelis, the “Father of Cats”.

    Joseph Leidy initially defined the genus Patriofelis in 1870 using fossils discovered in Middle Eocene soils in Wyoming; the type species is Patriofelis ulta, which was also discovered in Colorado. Fossils of P. ferox, another well-known species, have been discovered in somewhat older soils in Wyoming and Oregon.

    Patriofelis is typical of the Oxyaenodontas (“sharp tooth hyenas”), a widespread group of animals from the Paleocene and Eocene that exhibited extreme carnivory. Among the Oxyaenodonta, Patriofelis was both huge and highly specialized. The smaller Oxyaena, the related Malfelis genus, and the enormous Sarkastodon were its closest relatives.

    The Lifestyle of This Animal

    Patriofelis ferox by Charles R. Knight.
    Patriofelis ferox by Charles R. Knight.

    A Scavenger

    Despite its small size, Patriofelis was clearly an effective predator because of its robust set of teeth and smaller skull. Patriofelis must have been an active scavenger, since the skeleton seems to be designed to enable the animal to climb trees, despite this trait being more often associated with predators who feed on carrion. Not built for speed, this predator was better adapted to ambush hunting.

    Unique Adaptations

    Right hand of Patriofelis.
    Right hand of Patriofelis.

    Since the fossils of Patriofelis have been discovered at river deoposites and flood zones, this may imply that this animal was suited to walking on soft terrain, like riverbanks. Because of its small legs and widely spaced fingers, several researchers have hypothesized that Patriofelis was a strong swimmer. Given Patriofelis’ massive size, it’s hard to believe that this animal ever engaged in burrowing behavior, as suggested by the morphology of its upper arm bone (humerus) and hand.


    The Distinct Features of Patriofelis

    Powerful Jaw Muscles and a Strong Bite

    The cranium of this prehistoric animal was huge, measuring up to 10 inches (25 cm) in length and as big as a lion’s. It also had a short nose. In contrast to other creodonts (“meat teeth”), Patriofelis had some distinct skull features. For starters, it had special ear bones (ossified tympanic bulla) and passages in its skull that were likely linked to its hearing, sensory perception, facial expressions, etc.

    The depiction of the Patriofelis.
    The depiction of the Patriofelis.

    When we look at its skull, we see large ridges where muscles are attached, showing it had powerful jaw muscles. Its cheekbones were strong, and its lower jaw was sturdy despite being not very long. Interestingly, the teeth of this ancient animal were quite distinct. The upper premolars had sharp internal edges, and their incisors were smaller than their canines. It was also missing its top third molar. Additionally, the upper back teeth were shaped like sharp blades, indicating they likely had a strong bite.

    All these features give us clues about Patriofelis as a prehistoric mammal. It probably had a strong bite and might have been really good at hunting. Thanks to its ear bones and facial structures, it had acute senses like a modern felidae.


    A Shorter Body but a Muscular Tail

    Patriofelis had a somewhat sturdy skeleton. It had a shorter body and a neck of average length compared to other creodonts. The lumbar vertebrae in Patriofelis connected in a way that resembled how some animals with split hooves, like deer or cows, have their spines arranged.


    However, it was on a more complicated scale. A long, muscular tail curled out from the body towards the end.

    The lower extremities, and the forelegs of the Patriofelis in particular, were short and powerful. The shoulder blade of Patriofelis had specific features. It had two equal-sized depressions and large protrusions. Additionally, there was a visible bony projection. These shoulder blade characteristics indicate good mobility and advanced hunting techniques for Patriofelis. These features were generally typical among early meat-eating animals.

    Patriofelis shared similarities in its upper arm bone with its relative, the Oxyaena (“sharp hyena”), but it had more distinctive traits. Notably, it had a highly developed deltoid crest, which is a bony ridge where powerful shoulder muscles attach. Thus, Patriofelis likely had strong chest and shoulder muscles which makes it powerful as a predator.

    Powerful Leg Muscles

    As expected, the size of the olecranon (an elbow bone), along with other features, suggests that Patriofelis was exceptionally adept at walking. Its well-adapted limbs and muscles likely allowed it to move efficiently.

    Patriofelis also had a unique hip and leg structure. Its femur, the thigh bone, could move freely due to its long ilium (a part of the pelvic bone) which provided a sturdy base for attaching powerful gluteal muscles that are crucial for movement. A bony protrusion on the femur was positioned at a higher angle in Patriofelis which indicates robust leg muscles.

    Almost Dull Claws

    The limbs of this animal ended in pentadactyl hands and feet, meaning it had five digits on each. This is still a common trait among mammals after millions of years. However, Patriofelis had small and widely spaced toes with slightly blunt nail phalanges, suggesting that its claws weren’t extremely sharp.

    These features hint at its ability to walk effectively but it might not have been specialized for climbing or grasping prey with its claws because its claws probably weren’t that sharp.